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I too think it's from a burrowing animal. Instead of crab I would say it is some kind of lugworm/sandworm. I found this similar image: The keywords for this image contain "Wattwurm" = lugworm, though the image might be mislabeled.


This is almost exactly the opposite of what you think it is - it is actually from a burrowing animal, so instead of flying, it is actually burrowing into the sub-surface liquifaction layer of the sand and leaving behind this imprint. I think (correct me if I am wrong), but it is likely that the start of the burrow is the depression above your toe and the ...

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